49ers nose tackle Aubrayo Franklin ended his lengthy holdout Saturday by signing a $7.003 million franchise tag tender. San Francisco defensive coordinator Greg Manusky said this week that while Franklin had fallen behind his teammates by missing camp, he believed the nose tackle could get up to speed for the opener by playing in the exhibition finale next Thursday against San Diego. Franklin has recorded 291 tackles, four sacks, one interception and three forced fumbles in his seven-year career.